Raphanusraphanistrumandsativushaveoftenbeenrankedasdistinctspecies,andowingtodifferencesintheirfruitevenasdistinctgenera;
butProfessorHoffman(’Bot。Zeitung’1872page482)hasnowshownthatthesedifferences,remarkableastheyare,graduateaway,thefruitofR。
caudatusbeingintermediate。BycultivatingR。raphanistrumduringseveralgenerations(ibid1873page9),ProfessorHoffmanalsoobtainedplantsbearingfruitslikethoseofR。sativus。
PEA(Pisumsativum)。
Mostbotanistslookatthegarden—peaasspecificallydistinctfromthefield—pea(P。arvense)。ThelatterexistsinawildstateinSouthernEurope;buttheaboriginalparentofthegarden—peahasbeenfoundbyonecollectoralone,ashestates,intheCrimea。(9/81。Alph。DeCandolle’Geograph。Bot。’960。Mr。Bentham’Hort。Journal’volume91855page141
believesthatgardenandfieldpeasbelongtothesamespecies,andinthisrespecthediffersfromDr。Targioni。)AndrewKnightcrossed,asIaminformedbytheRev。A。Fitch,thefield—peawithawell—knowngardenvariety,thePrussianpea,andthecrossseemstohavebeenperfectlyfertile。Dr。Alefieldhasrecentlystudied(9/82。’BotanischeZeitung’1860
s。204。)thegenuswithcare,and,afterhavingcultivatedaboutfiftyvarieties,concludesthatcertainlytheyallbelongtothesamespecies。Itisaninterestingfactalreadyalludedto,that,accordingtoO。Heer(9/83。’DiePflanzenderPfahlbauten’1866s。23。)thepeasfoundinthelake—habitationsofSwitzerlandoftheStoneandBronzeages,belongtoanextinctvariety,withexceedinglysmallseeds,alliedtoP。arvenseorthefield—pea。Thevarietiesofthecommongarden—peaarenumerous,anddifferconsiderablyfromoneanother。ForcomparisonIplantedatthesametimeforty—one,EnglishandFrenchvarieties。Theydifferedgreatlyinheight,——
namelyfrombetween6and12inchesto8feet(9/84。AvarietycalledtheRouncivaattainsthisheight,asisstatedbyMr。Gordonin’Transact。
Hort。Soc。’2ndseriesvolume11835page374fromwhichpaperIhavetakensomefacts。),——inmannerofgrowth,andinperiodofmaturity。Somedifferingeneralaspectevenwhileonlytwoorthreeinchesinheight。ThestemsofthePrussianpeaaremuchbranched。Thetallkindshavelargerleavesthanthedwarfkinds,butnotinstrictproportiontotheirheight:——HAIR’S
DWARFMONMOUTHhasverylargeleaves,andthePOISNAINHATIF,andthemoderatelytallBLUEPRUSSIAN,haveleavesabouttwo—thirdsofthesizeofthetallestkind。IntheDANECROFTtheleafletsarerathersmallandalittlepointed;intheQUEENOFDWARFSratherrounded;andintheQUEENOF
ENGLANDbroadandlarge。Inthesethreepeastheslightdifferencesintheshapeoftheleavesareaccompaniedbyslightdifferencesincolour,inthePOISGEANTSANSPARCHEMIN,whichbearspurpleflowers,theleafletsintheyoungplantareedgedwithred;andinallthepeaswithpurpleflowersthestipulesaremarkedwithred。
Inthedifferentvarieties,one,two,orseveralflowersinasmallcluster,areborneonthesamepeduncle;andthisisadifferencewhichisconsideredofspecificvalueinsomeoftheLeguminosae。Inallthevarietiestheflowerscloselyresembleeachotherexceptincolourandsize。Theyaregenerallywhite,sometimespurple,butthecolourisinconstanteveninthesamevariety。InWARNER’SEMPEROR,whichisatallkind,theflowersarenearlydoublethesizeofthePOISNAINHATIF;butHAIR’SDWARFMONMOUTH,whichhaslargeleaves,likewisehaslargeflowers。
ThecalyxintheVICTORIAMARROWislarge,andinBISHOP’SLONGPODthesepalsarerathernarrow。Innootherkindisthereanydifferenceintheflower。
Thepodsandseeds,whichwithnaturalspeciesaffordsuchconstantcharacters,differgreatlyinthecultivatedvarietiesofthepea;andthesearethevaluable,andconsequentlytheselectedparts。SUGARPEAS,orPOISSANSPARCHEMIN,areremarkablefromtheirthinpods,which,whilstyoung,arecookedandeatenwhole;andinthisgroup,which,accordingtoMr。Gordonincludeselevensub—varieties,itisthepodwhichdiffersmost;
thusLEWIS’SNEGRO—PODDEDPEAhasastraight,broad,smooth,anddark—
purplepod,withthehusknotsothinasintheotherkinds;thepodofanothervarietyisextremelybowed;thatofthePOISGEANTismuchpointedattheextremity;andinthevariety"AGRANDSCOSSES"thepeasareseenthroughthehuskinsoconspicuousamannerthatthepod,especiallywhendry,canhardlyatfirstberecognisedasthatofapea。
Intheordinaryvarietiesthepodsalsodiffermuchinsize;——incolour,thatofWOODFORD’SGREENMARROWbeingbright—greenwhendry,insteadofpalebrown,andthatofthepurple—poddedpeabeingexpressedbyitsname;—
—insmoothness,thatofDANECROFTbeingremarkablyglossy,whereasthatoftheNEPLUSULTRAisrugged;inbeingeithernearlycylindrical,orbroadandflat;——inbeingpointedattheend,asinTHURSTON’SRELIANCE,ormuchtruncated,asintheAMERICANDWARF。IntheAUVERGNEPEAthewholeendofthepodisbowedupwards。IntheQUEENOFTHEDWARFSandinSCIMITARPEAS
thepodisalmostellipticinshape。Iheregivedrawingsofthefourmostdistinctpodsproducedbytheplantscultivatedbyme。
(FIGURE41。PODSANDPEAS。I。QueenofDwarfs。II。AmericanDwarf。III。
Thurston’sReliance。IV。PoisGeantsansparchemin。a。DanO’RourkePea。b。
QueenofDwarfsPea。c。Knight’sTallWhiteMarrow。a。Lewis’sNegroPea。)
Inthepeaitselfwehaveeverytintbetweenalmostpurewhite,brown,yellow,andintensegreen;inthevarietiesoftheSUGARPEASwehavethesesametints,togetherwithredpassingthroughfinepurpleintoadarkchocolatetint。Thesecoloursareeitheruniformordistributedindots,striae,ormoss—likemarks;theydependinsomecasesonthecolourofthecotyledonsseenthroughtheskin,andinothercasesontheoutercoatsofthepeaitself。Inthedifferentvarieties,thepodscontain,accordingtoMr。Gordon,fromelevenortwelvetoonlyfourorfivepeas。Thelargestpeasarenearlytwiceasmuchindiameterasthesmallest;andthelatterarenotalwaysbornebythemostdwarfedkinds。Peasdiffermuchinshape,beingsmoothandspherical,smoothandoblong,nearlyovalintheQUEENOF
THEDWARFS,andnearlycubicalandcrumpledinmanyofthelargerkinds。
Withrespecttothevalueofthedifferencesbetweenthechiefvarieties,itcannotbedoubtedthat,ifoneofthetallSUGAR—PEAS,withpurpleflowers,thin—skinnedpodsofanextraordinaryshape,includinglarge,dark—purplepeas,grewwildbythesideofthelowlyQUEENOFTHEDWARFS,withwhiteflowers,greyish—green,roundedleaves,scimitar—likepods,containingoblong,smooth,pale—colouredpeas,whichbecamematureatadifferentseason:orbythesideofoneofthegiganticsorts,liketheCHAMPIONOFENGLAND,withleavesofgreatsize,pointedpods,andlarge,green,crumpled,almostcubicalpeas,——allthreekindswouldberankedasdistinctspecies。
AndrewKnight(9/85。’Phil。Tract。’1799page196。)hasobservedthatthevarietiesofpeaskeepverytrue,becausetheyarenotcrossedbyinsects。
Asfarasthefactofkeepingtrueisconcerned,IhearfromMr。MastersofCanterbury,wellknownastheoriginatorofseveralnewkinds,thatcertainvarietieshaveremainedconstantforaconsiderabletime,——forinstance,KNIGHT’SBLUEDWARF,whichcameoutabouttheyear1820。(9/86。’Gardener’sMagazine’volume11826page153。)Butthegreaternumberofvarietieshaveasingularlyshortexistence:thusLoudonremarks(9/87。’EncyclopaediaofGardening’page823。)that"sortswhichwerehighlyapprovedin1821,arenow,in1833,nowheretobefound;"andoncomparingthelistsof1833withthoseof1855,Ifindthatnearlyallthevarietieshavechanged。Mr。
Mastersinformsmethatthenatureofthesoilcausessomevarietiestolosetheircharacter。Aswithotherplants,certainvarietiescanbepropagatedtruly,whilstothersshowadeterminedtendencytovary;thustwopeasdifferinginshape,oneroundandtheotherwrinkled,werefoundbyMr。Masterswithinthesamepod,buttheplantsraisedfromthewrinkledkindalwaysevincedastrongtendencytoproduceroundpeas。Mr。Mastersalsoraisedfromaplantofanothervarietyfourdistinctsub—varieties,whichboreblueandround,whiteandround,blueandwrinkled,andwhiteandwrinkledpeas;andalthoughhesowedthesefourvarietiesseparatelyduringseveralsuccessiveyears,eachkindalwaysreproducedallfourkindsmixedtogether!
Withrespecttothevarietiesnotnaturallyintercrossing,Ihaveascertainedthatthepea,whichinthisrespectdiffersfromsomeotherLeguminosae,isperfectlyfertilewithouttheaidofinsects。YetIhaveseenhumble—beeswhilstsuckingthenectardepressthekeel—petals,andbecomesothicklydustedwithpollen,thatitcouldhardlyfailtobeleftonthestigmaofthenextflowerwhichwasvisited。Nevertheless,distinctvarietiesgrowingcloselytogetherrarelycross;andIhavereasontobelievethatthisisduetotheirstigmasbeingprematurelyfertilisedinthiscountrybypollenfromthesameflower。Thehorticulturistswhoraiseseed—peasarethusenabledtoplantdistinctvarietiesclosetogetherwithoutanybadconsequences;anditiscertain,asIhavemyselffound,thattrueseedmaybesavedduringatleastseveralgenerationsunderthesecircumstances。(9/88。SeeDr。Andersontothesameeffectinthe’BathSoc。
AgriculturalPapers’volume4page87。)Mr。Fitchraised,asheinformsme,onevarietyfortwentyyears,anditalwayscametrue,thoughgrownclosetoothervarieties。Fromtheanalogyofkidney—beansIshouldhaveexpected(9/89。Ihavepublishedfulldetailsofexperimentsonthissubjectinthe’Gardener’sChronicle’1857October25。)thatvarietiesthuscircumstancedwouldhaveoccasionallycrossed;andIshallgiveintheeleventhchaptertwocasesofthishavingoccurred,asshown(inamannerhereaftertobeexplained)bythepollenoftheonevarietyhavingacteddirectlyontheseedsoftheother。Whethermanyofthenewvarietieswhichincessantlyappearareduetosuchoccasionalandaccidentalcrosses,Idonotknow。
NordoIknowwhethertheshortexistenceofalmostallthenumerousvarietiesistheresultofmerechangeoffashion,oroftheirhavingaweakconstitution,frombeingtheproductoflong—continuedself—
fertilisation。Itmay,however,benoticedthatseveralofAndrewKnight’svarieties,whichhaveenduredlongerthanmostkinds,wereraisedtowardsthecloseofthelastcenturybyartificialcrosses;someofthem,I
believe,werestillvigorousin1860;butnow,in1865,awriter,speaking(9/90。’Gardener’sChronicle’1865page387。)ofKnight’sfourkindsofmarrows,says,theyhaveacquiredafamoushistory,buttheirgloryhasdeparted。
WithrespecttoBeans(Fabavulgaris),Iwillsaybutlittle。Dr。Alefieldhasgiven(9/91。’Bonplandia’10,1862s。348。)shortdiagnosticcharactersoffortyvarieties。Everyonewhohasseenacollectionmusthavebeenstruckwiththegreatdifferenceinshape,thickness,proportionallengthandbreadth,colour,andsizewhichbeanspresent。WhatacontrastbetweenaWindsorandHorse—bean!Asinthecaseofthepea,ourexistingvarietieswereprecededduringtheBronzeageinSwitzerland(9/92。Heer’DiePflanzenderPfahlbauten’1866s。22。)byapeculiarandnowextinctvarietyproducingverysmallbeans。(9/93。Mr。BenthaminformsmethatinPoitouandtheadjoiningpartsofFrance,varietiesofPhaseolusvulgarisareextremelynumerous,andsodifferentthattheyweredescribedbySaviasdistinctspecies。Mr。Benthambelievesthatallaredescendedfromanunknowneasternspecies。Althoughthevarietiesdiffersogreatlyinstatureandintheirseeds,"thereisaremarkablesamenessintheneglectedcharactersoffoliageandflowers,andespeciallyinthebracteoles,aninsignificantcharacterintheeyesevenofbotanists。")
POTATO(Solanumtuberosum)。
Thereislittledoubtabouttheparentageofthisplant;forthecultivatedvarietiesdifferextremelylittleingeneralappearancefromthewildspecies,whichcanberecognisedinitsnativelandatthefirstglance。
(9/94。Darwin’JournalofResearches’1845page285。Sabinein’Transact。
Hort。Soc。’volume5page249。)ThevarietiescultivatedinBritainarenumerous;thusLawson(9/95。’SynopsisoftheVegetableProductsofScotland’quotedinWilson’s’BritishFarming’page317。)givesadescriptionof175kinds。Iplantedeighteenkindsinadjoiningrows;theirstemsandleavesdifferedbutlittle,andinseveralcasestherewasasgreatadifferencebetweentheindividualsofthesamevarietyasbetweenthedifferentvarieties。Theflowervariedinsize,andincolourbetweenwhiteandpurple,butinnootherrespect,exceptthatinonekindthesepalsweresomewhatelongated。Onestrangevarietyhasbeendescribedwhichalwaysproducestwosortsofflowers,thefirstdoubleandsterile,thesecondsingleandfertile。(9/96。SirG。Mackenzie,in’Gardener’sChronicle’1845page790。)Thefruitorberriesalsodiffer,butonlyinaslightdegree。(9/97。PutscheundVertuch’VersucheinerMonographiederKartoffeln’1819s。9,15。SeealsoDr。Anderson’RecreationsinAgriculture’volume4page325。)ThevarietiesareliableinverydifferentdegreetotheattackoftheColoradopotato—beetle。(9/98。Walsh’TheAmericanEntomologist’1869page160。AlsoS。Tenney’TheAmericanNaturalist’May1871page171。)
Thetubers,ontheotherhand,presentawonderfulamountofdiversity。
Thisfactaccordswiththeprinciplethatthevaluableandselectedpartsofallcultivatedproductionspresentthegreatestamountofmodification。
Theydiffermuchinsizeandshape,beingglobular,oval,flattened,kidney—like,orcylindrical。OnevarietyfromPeruisdescribed(9/99。
’Gardener’sChronicle’1862page1052。)asbeingquitestraight,andatleastsixinchesinlength,thoughnothickerthanaman’sfinger。Theeyesorbudsdifferinform,position,andcolour。Themannerinwhichthetubersarearrangedontheso—calledrootsorrhizomesisdifferent;thus,inthegurken—kartoffelntheyformapyramidwiththeapexdownwards,andinanothervarietytheyburythemselvesdeepintheground。Therootsthemselvesruneithernearthesurfaceordeepintheground。Thetubersalsodifferinsmoothnessandcolour,beingexternallywhite,red,purple,oralmostblack,andinternallywhite,yellow,oralmostblack。Theydifferinflavourandquality,beingeitherwaxyormealy;intheirperiodofmaturity,andintheircapacityforlongpreservation。
Aswithmanyotherplantswhichhavebeenlongpropagatedbybulbs,tubers,cuttings,etc。,bywhichmeansthesameindividualisexposedduringale...完整阅读请扫描二维码下载丁香书院APP免费看: