首页 >出版文学> The Variation of Animals and Plants under Domestic>第36章
  Indomitablepatience,thefinestpowersofdiscrimination,andsoundjudgmentmustbeexercisedduringmanyyears。Aclearlypredeterminedobjectmustbekeptsteadilyinview。Fewmenareendowedwithallthesequalities,especiallywiththatofdiscriminatingveryslightdifferences;judgmentcanbeacquiredonlybylongexperience;butifanyofthesequalitiesbewanting,thelabourofalifemaybethrownaway。Ihavebeenastonishedwhencelebratedbreeders,whoseskillandjudgmenthavebeenprovedbytheirsuccessatexhibitions,haveshownmetheiranimals,whichappearedallalike,andhaveassignedtheirreasonsformatchingthisandthatindividual。TheimportanceofthegreatprincipleofSelectionmainlyliesinthispowerofselectingscarcelyappreciabledifferences,whichneverthelessarefoundtobetransmissible,andwhichcanbeaccumulateduntiltheresultismademanifesttotheeyesofeverybeholder。
  Theprincipleofselectionmaybeconvenientlydividedintothreekinds。
  METHODICALSELECTIONisthatwhichguidesamanwhosystematicallyendeavourstomodifyabreedaccordingtosomepredeterminedstandard。UNCONSCIOUS
  SELECTIONisthatwhichfollowsfrommennaturallypreservingthemostvaluedanddestroyingthelessvaluedindividuals,withoutanythoughtofalteringthebreed;andundoubtedlythisprocessslowlyworksgreatchanges。
  Unconsciousselectiongraduatesintomethodical,andonlyextremecasescanbedistinctlyseparated;forhewhopreservesausefulorperfectanimalwillgenerallybreedfromitwiththehopeofgettingoffspringofthesamecharacter;butaslongashehasnotapredeterminedpurposetoimprovethebreed,hemaybesaidtobeselectingunconsciously。(20/1。Theterm"unconsciousselection"hasbeenobjectedtoasacontradiction;butseesomeexcellentobservationsonthisheadbyProf。Huxley(’Nat。Hist。Review’
  October1864page578)whoremarksthatwhenthewindheapsupsand—dunesitsiftsandUNCONSCIOUSLYSELECTSfromthegravelonthebeachgrainsofsandofequalsize。)Lastly,wehaveNATURALSELECTION,whichimpliesthattheindividualswhicharebestfittedforthecomplex,andinthecourseofageschangingconditionstowhichtheyareexposed,generallysurviveandprocreatetheirkind。Withdomesticproductions,naturalselectioncomestoacertainextentintoaction,independentlyof,andeveninoppositionto,thewillofman。
  METHODICALSELECTION。
  WhatmanhaseffectedwithinrecenttimesinEnglandbymethodicalselectionisclearlyshownbyourexhibitionsofimprovedquadrupedsandfancybirds。
  Withrespecttocattle,sheep,andpigs,weowetheirgreatimprovementtoalongseriesofwell—knownnames——Bakewell,Coiling,Ellman,Bates,JonasWebb,LordsLeicesterandWestern,FisherHobbs,andothers。Agriculturalwritersareunanimousonthepowerofselection:anynumberofstatementstothiseffectcouldbequoted;afewwillsuffice。Youatt,asagaciousandexperiencedobserver,writes(20/2。’OnSheep’1838page60。)theprincipleofselectionis"thatwhichenablestheagriculturist,notonlytomodifythecharacterofhisflock,buttochangeitaltogether。"AgreatbreederofShorthorns(20/3。Mr。J。WrightonShorthornCattlein’JournalofRoyalAgricult。Soc。’volume7pages208,209。)says,"IntheanatomyoftheshouldermodernbreedershavemadegreatimprovementontheKettonshorthornsbycorrectingthedefectintheknuckleorshoulder—joint,andbylayingthetopoftheshouldermoresnuglyinthecrop,andtherebyfillingupthehollowbehindit……Theeyehasitsfashionatdifferentperiods:atonetimetheeyehighandoutstandingfromthehead,andatanothertimethesleepyeyesunkintothehead;buttheseextremeshavemergedintothemediumofafull,clearandprominenteyewithaplacidlook。"
  Again,hearwhatanexcellentjudgeofpigs(20/4。H。D。Richardson’OnPigs’
  1847page44。)says:"Thelegsshouldbenolongerthanjusttopreventtheanimal’sbellyfromtrailingontheground。Thelegistheleastprofitableportionofthehog,andwethereforerequirenomoreofitthanisabsolutelynecessaryforthesupportoftherest。"Letanyonecomparethewild—boarwithanyimprovedbreed,andhewillseehoweffectuallythelegshavebeenshortened。
  Fewpersons,exceptbreeders,areawareofthesystematiccaretakeninselectinganimals,andofthenecessityofhavingaclearandalmostpropheticvisionintofuturity。LordSpencer’sskillandjudgmentwerewellknown;andhewrites(20/5。’JournalofRoyalAgricult。Soc。’volume1page24。),"Itisthereforeverydesirable,beforeanymancommencestobreedeithercattleorsheep,thatheshouldmakeuphismindtotheshapeandqualitieshewishestoobtain,andsteadilypursuethisobject。"LordSomerville,inspeakingofthemarvellousimprovementoftheNewLeicestersheep,effectedbyBakewellandhissuccessors,says,"Itwouldseemasiftheyhadfirstdrawnaperfectform,andthengivenitlife。"Youatt(20/6。’OnSheep’pages520,319。)urgesthenecessityofannuallydraftingeachflock,asmanyanimalswillcertainlydegenerate"fromthestandardofexcellencewhichthebreederhasestablishedinhisownmind。"Evenwithabirdofsuchlittleimportanceasthecanary,longago(1780—1790)ruleswereestablished,andastandardofperfectionwasfixedaccordingtowhichtheLondonfancierstriedtobreedtheseveralsub—
  varieties。(20/7。Loudon’s’Mag。ofNat。Hist。’volume81835page618。)A
  greatwinnerofprizesatthePigeon—shows(20/8。’AtreatiseontheArtofBreedingtheAlmondTumbler’1851page9。),indescribingtheshort—facedAlmondTumbler,says,"Therearemanyfirst—ratefancierswhoareparticularlypartialtowhatiscalledthegoldfinch—beak,whichisverybeautiful;otherssay,takeafull—sizeroundcherrythentakeabarleycorn,andjudiciouslyplacingandthrustingitintothecherry,formasitwereyourbeak;andthatisnotall,foritwillformagoodheadandbeak,provided,asIsaidbefore,itisjudiciouslydone;otherstakeanoat;butasIthinkthegoldfinch—beakthehandsomest,Iwouldadvisetheinexperiencedfanciertogettheheadofagoldfinch,andkeepitbyhimforhisobservation。"Wonderfullydifferentasarethebeaksoftherockpigeonandgoldfinch,theendhasundoubtedlybeennearlygained,asfarasexternalshapeandproportionsareconcerned。
  Notonlyshouldouranimalsbeexaminedwiththegreatestcarewhilstalive,but,asAndersonremarks(20/9。’RecreationsinAgriculture’volume2page409。)theircarcasesshouldbescrutinised,"soastobreedfromthedescendantsofsuchonlyas,inthelanguageofthebutcher,cutupwell。"The"grainofthemeat"incattle,anditsbeingwellmarbledwithfat(20/10。
  ’YouattonCattle’pages191,227。),andthegreaterorlessaccumulationoffatintheabdomenofoursheep,havebeenattendedtowithsuccess。Sowithpoultry,awriter(20/11。Ferguson’PrizePoultry’1854page208。),speakingofCochin—Chinafowls,whicharesaidtodiffermuchinthequalityoftheirflesh,says,"thebestmodeistopurchasetwoyoungbrother—cocks,kill,dress,andserveupone;ifhebeindifferent,similarlydisposeoftheother,andtryagain;if,however,hebefineandwell—flavoured,hisbrotherwillnotbeamissforbreedingpurposesforthetable。"
  Thegreatprincipleofthedivisionoflabourhasbeenbroughttobearonselection。Incertaindistricts(20/12。Wilsonin’Transact。HighlandAgricult。Soc。’quotedin’Gardener’sChronicle’1844page29。)"thebreedingofbullsisconfinedtoaverylimitednumberofpersons,whobydevotingtheirwholeattentiontothisdepartment,areablefromyeartoyeartofurnishaclassofbullswhicharesteadilyimprovingthegeneralbreedofthedistrict。"Therearingandlettingofchoiceramshaslongbeen,asiswellknown,achiefsourceofprofittoseveraleminentbreeders。InpartsofGermanythisprincipleiscarriedwithmerinosheeptoanextremepoint。
  (20/13。Simmondsquotedin’Gardener’sChronicle’1855page637。Andforthesecondquotationsee’YouattonSheep’page171。)So"importantistheproperselectionofbreedinganimalsconsidered,thatthebestflock—mastersdonottrusttotheirownjudgmentortothatoftheirshepherds,butemploypersonscalled’sheep—classifiers’whomakeittheirspecialbusinesstoattendtothispartofthemanagementofseveralflocks,andthustopreserve,orifpossibletoimprove,thebestqualitiesofbothparentsinthelambs。"InSaxony,"whenthelambsareweaned,eachinhisturnisplaceduponatablethathiswoolandformmaybeminutelyobserved。Thefinestareselectedforbreedingandreceiveafirstmark。Whentheyareoneyearold,andpriortoshearingthem,anothercloseexaminationofthosepreviouslymarkedtakesplace:thoseinwhichnodefectcanbefoundreceiveasecondmark,andtherestarecondemned。Afewmonthsafterwardsathirdandlastscrutinyismade;
  theprimeramsandewesreceiveathirdandfinalmark,buttheslightestblemishissufficienttocausetherejectionoftheanimal。"Thesesheeparebredandvaluedalmostexclusivelyforthefinenessoftheirwool;andtheresultcorrespondswiththelabourbestowedontheirselection。Instrumentshavebeeninventedtomeasureaccuratelythethicknessofthefibres;and"anAustrianfleecehasbeenproducedofwhichtwelvehairsequalledinthicknessonefromaLeicestersheep。"
  Throughouttheworld,whereversilkisproduced,thegreatestcareisbestowedonselectingthecocoonsfromwhichthemothsforbreedingaretobereared。A
  carefulcultivator(20/14。Robinet’VersaSoie’1848page271。)likewiseexaminesthemothsthemselves,anddestroysthosethatarenotperfect。ButwhatmoreimmediatelyconcernsusisthatcertainfamiliesinFrancedevotethemselvestoraisingeggsforsale。(20/15。Quatrefages’LesMaladiesduVeraSoie’1859page101。)InChina,nearShanghai,theinhabitantsoftwosmalldistrictshavetheprivilegeofraisingeggsforthewholesurroundingcountry,andthattheymaygiveuptheirwholetimetothisbusiness,theyareinterdictedbylawfromproducingsilk。(20/16。M。Simonin’Bull。delaSoc。
  d’Acclimat。’tome91862page221。)
  Thecarewhichsuccessfulbreederstakeinmatchingtheirbirdsissurprising。
  SirJohnSebright,whosefameisperpetuatedbythe"SebrightBantam,"usedtospend"twoandthreedaysinexamining,consulting,anddisputingwithafriendwhichwerethebestoffiveorsixbirds。"(20/17。’ThePoultryChronicle’volume11854page607。)Mr。Bult,whosepouter—pigeonswonsomanyprizes,andwereexportedtoNorthAmericaunderthechargeofamansentonpurpose,toldmethathealwaysdeliberatedforseveraldaysbeforehematchedeachpair。Hencewecanunderstandtheadviceofaneminentfancier,whowrites(20/18。J。M。Eaton’ATreatiseonFancyPigeons’1852page14and’A
  TreatiseontheAlmondTumbler’1851page11。)"Iwouldhereparticularlyguardyouagainsthavingtoogreatavarietyofpigeons,otherwiseyouwillknowalittleofall,butnothingaboutoneasitoughttobeknown。"
  Apparentlyittranscendsthepowerofthehumanintellecttobreedallkinds:
  "itispossiblethattheremaybeafewfanciersthathaveagoodgeneralknowledgeoffancypigeons;buttherearemanymorewholabourunderthedelusionofsupposingtheyknowwhattheydonot。"Theexcellenceofonesub—
  variety,theAlmondTumbler,liesintheplumage,carriage,head,beak,andeye;butitistoopresumptuousinthebeginnertotryforallthesepoints。
  Thegreatjudgeabovequotedsays,"Therearesomeyoungfancierswhoareover—covetous,whogoforalltheabovefivepropertiesatonce;theyhavetheirrewardbygettingnothing。"Wethusseethatbreedingevenfancypigeonsisnosimpleart:wemaysmileatthesolemnityoftheseprecepts,buthewholaughswillwinnoprizes。
  Whatmethodicalselectionhaseffectedforouranimalsissufficientlyproved,asalreadyremarked,byourExhibitions。Sogreatlywerethesheepbelongingtosomeoftheearlierbreeders,suchasBakewellandLordWestern,changed,thatmanypersonscouldnotbepersuadedthattheyhadnotbeencrossed。Ourpigs,asMr。Corringhamremarks(20/19。’JournalRoyalAgriculturalSoc。’
  volume6page22。)duringthelasttwentyyearshaveundergone,throughrigorousselectiontogetherwithcrossing,acompletemetamorphosis。ThefirstexhibitionforpoultrywasheldintheZoologicalGardensin1845;andtheimprovementeffectedsincethattimehasbeengreat。AsMr。Bailey,thegreatjudge,remarkedtome,itwasformerlyorderedthatthecomboftheSpanishcockshouldbeupright,andinfourorfiveyearsallgoodbirdshaduprightcombs;itwasorderedthatthePolishcockshouldhavenocomborwattles,andnowabirdthusfurnishedwouldbeatoncedisqualified;beardswereordered,andoutoffifty—sevenpenslately(1860)exhibitedattheCrystalPalace,allhadbeards。Soithasbeeninmanyothercases。Butinallcasesthejudgesorderonlywhatisoccasionallyproducedandwhatcanbeimprovedandrenderedconstantbyselection。Thesteadyincreaseinweightduringthelastfewyearsinourfowls,turkeys,ducks,andgeeseisnotorious;"six—poundducksarenowcommon,whereasfourpoundswasformerlytheaverage。"Asthetimerequiredtomakeachangehasnotoftenbeenrecorded,itmaybeworthmentioningthatittookMr。Wickingthirteenyearstoputacleanwhiteheadonanalmondtumbler’sbody,"atriumph,"saysanotherfancier,"ofwhichhemaybejustlyproud。"(20/20。’PoultryChronicle’volume21855page596。)
  Mr。Tollet,ofBetleyHall,selectedcows,andespeciallybulls,descendedfromgoodmilkers,forthesolepurposeofimprovinghiscattlefortheproductionofcheese;hesteadilytestedthemilkwiththelactometer,andineightyearsheincreased,asIwasinformedbyhim,theproductinproportionoffourtothree。Hereisacuriouscase(20/21。Isid。GeoffroySt。—Hilaire’Hist。Nat。Gen。’tome3page254。)ofsteadybutslowprogress,withtheendnotasyetfullyattained:in1784araceofsilkwormswasintroducedintoFrance,inwhichonehundredinthethousandfailedtoproducewhitecocoons;
  butnowaftercarefulselectionduringsixty—fivegenerations,theproportionofyellowcocoonshasbeenreducedtothirty—fiveinthethousand。
  Withplantsselectionhasbeenfollowedwiththesamegoodresultaswithanimals。Buttheprocessissimpler,forplantsinthegreatmajorityofcasesbearbothsexes。Nevertheless,withmostkindsitisnecessarytotakeasmuchcaretopreventcrossesaswithanimalsorunisexualplants;butwithsomeplants,suchaspeas,thiscareisnotnecessary。Withallimprovedplants,exceptingofcoursethosewhicharepropagatedbybuds,cuttings,etc。,itisalmostindispensabletoexaminetheseedlingsanddestroythosewhichdepartfromthepropertype。Thisiscalled"roguing,"andis,infact,aformofselection,liketherejectionofinferioranimals。Experiencedhorticulturistsandagriculturistsincessantlyurgeeveryonetopreservethefinestplantsfortheproductionofseed。
  Althoughplantsoftenpresentmuchmoreconspicuousvariationsthananimals,yettheclosestattentionisgenerallyrequisitetodetecteachslightandfavourablechange。Mr。Mastersrelates(20/22。’Gardener’sChronicle’1850
  page198。)how"manyapatienthourwasdevoted,"whilsthewasyoung,tothedetectionofdifferencesinpeasintendedforseed。Mr。Barnet(20/23。
  ’Transact。Hort。Soc。’volume6page152。)remarksthattheoldscarletAmericanstrawberrywascultivatedformorethanacenturywithoutproducingasinglevariety;andanotherwriterobserveshowsingularitwasthatwhengardenersfirstbegantoattendtothisfruititbegantovary;thetruthnodoubtbeingthatithadalwaysvaried,butthat,untilslightvariationswereselectedandpropagatedbyseed,noconspicuousresultwasobtained。Thefinestshadesofdifferenceinwheathavebeendiscriminatedandselectedwithalmostasmuchcareas,inthecaseofthehigheranimals,forinstancebyCol。LeCouteurandmoreespeciallybyMajorHallett。
  Itmaybeworthwhiletogiveafewexamplesofmethodicalselectionwithplants;butinfactthegreatimprovementofallourancientlycul...完整阅读请扫描二维码下载丁香书院APP免费看

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