Thatisperfectlytrue;butarethereanyalterations?Tobeginwith,itseemstomehardlypossiblethattheinsectcangobacktothecuttingoncethepieceisdetachedfromtheleaf:itlacksthenecessarysupporttognawtheflimsydiskwithanyprecision。A
tailorwouldspoilhisclothifhehadnotthesupportofatablewhencuttingoutthepiecesforacoat。TheMegachile'sscissors,sodifficulttowieldonanythingnotfirmlyheld,woulddoequallybadwork。
Besides,IhavebetterevidencethanthisformyrefusaltobelieveintheexistenceofalterationswhentheBeehasthecellinfrontofher。Thelidiscomposedofapileofdiskswhosenumbersometimesreacheshalfascore。Nowthebottompartofallthesedisksistheundersurfaceoftheleaf,whichispalerandmorestronglyveined;
thetoppartistheuppersurface,whichissmoothandgreener。Inotherwords,theinsectplacestheminthepositionwhichtheyoccupywhengathered。Letmeexplain。Inordertocutoutapiece,theBeestandsontheuppersurfaceoftheleaf。Thepiecedetachedisheldinthefeetandisthereforelaidwithitstopsurfaceagainsttheinsect'schestatthemomentofdeparture。Thereisnopossibilityofitsbeingturnedoveronthejourney。Consequently,thepieceislaidastheBeehasjustpickedit,withthelowersurfacetowardstheinsideofthecellandtheuppersurfacetowardstheoutside。Ifalterationswerenecessarytoreducethelidtothediameterofthepot,thediskwouldbeboundtogetturnedover:thepiece,manipulated,setupright,turnedround,triedthiswayandthat,would,whenfinallylaidinposition,haveitstoporbottomsurfaceinsidejustasithappenedtocome。Butthisisexactlywhatdoesnottakeplace。Therefore,astheorderofstackingneverchanges,thedisksarecut,fromthefirstclipofthescissors,withtheirproperdimensions。Theinsectexcelsusinpracticalgeometry。IlookupontheLeaf—cutter'spotandlidasanadditiontothemanyothermarvelsofinstinctthatcannotbeexplainedbymechanics;Isubmitittotheconsiderationofscience;andIpasson。
TheSilkyLeaf—cutter(Megachilesericans,FONSCOL。;M。Dufourii,LEP。)makeshernestsinthedisusedgalleriesoftheAnthophorae。I
knowhertooccupyanotherdwellingwhichismoreelegantandaffordsamoreroomyinstallation:ImeantheolddwellingofthefatCapricorn,thedenizenoftheoaks。Themetamorphosisiseffectedinaspaciouschamberlinedwithsoftfelt。Whenthelong—hornedBeetlereachestheadultstage,hereleaseshimselfandemergesfromthetreebyfollowingavestibulewhichthelarva'spowerfultoolshavepreparedbeforehand。Whenthedesertedcabin,owingtoitsposition,remainswholesomeandthereisnosignofanyrunningfromitswalls,nobrownstuffsmellingofthetan—yard,itissoonvisitedbytheSilkyMegachile,whofindsinitthemostsumptuousoftheapartmentsinhabitedbytheLeaf—cutters。Itcombineseveryconditionofcomfort:perfectsafety,aneventemperature,freedomfromdamp,ampleroom;andsothemotherwhoisfortunateenoughtobecomethepossessorofsuchalodgingusesitentirely,vestibuleanddrawing—
roomalike。Accommodationisfoundforallherfamilyofeggs;atleast,Ihavenowhereseennestsaspopulousashere。
Oneofthemprovidesmewithseventeencells,thehighestnumberappearinginmycensusoftheMegachileclan。MostofthemarelodgedinthenymphalchamberoftheCapricorn;and,asthespaciousrecessistoowideforasinglerow,thecellsarearrangedinthreeparallelseries。Theremainder,inasinglestring,occupythevestibule,whichiscompletedandfilledupbytheterminalbarricade。Inthematerialsemployed,hawthorn—andpaliurus—leavespredominate。Thepieces,bothinthecellsandinthebarrier,varyinsize。Itistruethatthehawthorn—leaves,withtheirdeepindentations,donotlendthemselvestothecuttingofneatovalpieces。Theinsectseemstohavedetachedeachmorselwithouttroublingovermuchabouttheshapeofthepiece,solongasitwasbigenough。Norhasitbeenveryparticularaboutarrangingthepiecesaccordingtothenatureoftheleaf:afterafewbitsofpaliuruscomebitsofvineandhawthorn;andtheseagainarefollowedbybitsofbrambleandpaliurus。TheBeehascollectedherpiecesanyhow,takingabithereandthere,justasherfancydictated。
Nevertheless,paliurusisthecommonest,perhapsforeconomicalreasons。
Inotice,infact,thattheleavesofthisshrub,insteadofbeingusedpiecemeal,areemployedwhole,whentheydonotexceedtheproperdimensions。Theirovalformandtheirmoderatesizesuittheinsect'srequirements;andthereisthereforenonecessitytocutthemintopieces。Theleaf—stalkisclippedwiththescissorsand,withoutmoreado,theMegachileretiresthericherbyafirst—ratebitofmaterial。
Splitupintotheircomponentparts,twocellsgivemealtogethereighty—threepiecesofleaves,whereofeighteenaresmallerthantheothersandofaroundshape。Thelast—namedcomefromthelids。Iftheyaverageforty—twoeach,theseventeencellsofthenestrepresentsevenhundredandfourteenpieces。Thesearenotall:thenestends,intheCapricorn'svestibule,withastoutbarricadeinwhichIcountthreehundredandfiftypieces。Thetotalthereforeamountstoonethousandandsixty—four。AllthosejourneysandallthatworkwiththescissorstofurnishthedesertedchamberoftheCerambyx!IfIdidnotknowtheLeaf—cutter'ssolitaryandjealousdisposition,Ishouldattributethehugestructuretothecollaborationofseveralmothers;butthereisnoquestionofcommunisminthiscase。Onedauntlesscreatureandonealone,onesolitary,inveterateworker,hasproducedthewholeoftheprodigiousmass。Ifworkisthebestwaytoenjoylife,thisonecertainlyhasnotbeenboredduringthefewweeksofherexistence。
Igladlyawardherthemosthonourableofeulogies,thatduetotheindustrious;andIalsocomplimentheronhertalentforclosingthehoney—pots。Thepiecesstackedintolidsareroundandhavenothingtosuggestthoseofwhichthecellsandthefinalbarricadearemade。
Exceptingthefirst,thosenearestthehoney,theyareperhapscutalittlelessneatlythanthedisksoftheWhite—girdledLeaf—cutter;
nomatter:theystopthejarperfectly,especiallywhentherearesometenofthemoneabovetheother。Whencuttingthem,theBeewasassureofherscissorsasadressmakerguidedbyapatternlaidonthestuff;andyetshewascuttingwithoutamodel,withouthavinginfrontofherthemouthtobeclosed。Toenlargeonthisinterestingsubjectwouldmeantorepeatoneself。AlltheLeaf—cuttershavethesametalentformakingthelidsoftheirpots。
Alessmysteriousquestionthanthisgeometricalproblemisthatofthematerials。DoeseachspeciesofMegachilekeeptoasingleplant,orhasitadefinitebotanicaldomainwhereintoexerciseitslibertyofchoice?ThelittlethatIhavealreadysaidisenoughtomakeussuspectthattheinsectisnotrestrictedtooneplant;andthisisconfirmedbyanexaminationoftheseparatecells,piecebypiece,whenwefindavarietywhichwewerefarfromimaginingatfirst。
HereisthefloraoftheMegachilesinmyneighbourhood,averyincompletefloraanddoubtlesscapableofconsiderableamplificationbyfutureresearches。
TheSilkyLeaf—cuttergathersthematerialsforherpots,herlidsandherbarricadesfromthefollowingplants:paliurus,hawthorn,vine,wildbriar,bramble,holm—oak,amelanchier,terebinthus,sage—
leavedrock—rose。Thefirstthreesupplythegreaterpartoftheleaf—work;thelastthreearerepresentedonlybyrarefragments。
TheHare—footedLeaf—cutter(Megachilelagopoda,LIN。)whichIseeverybusyinmyenclosure,thoughsheonlycollectshermaterialsthere,exploitsthelilacandtherose—treebypreference。Fromtimetotime,Iseeheralsocuttingbitsoutoftherobinia,thequince—
treeandthecherry—tree。Intheopencountry,Ihavefoundherbuildingwiththeleavesofthevinealone。
TheSilveryLeaf—cutter(Megachileargentata,FAB。),anotherofmyguests,sharesthetasteoftheaforesaidforthelilacandtherose,butherdomainincludesinadditionthepomegranate—tree,thebramble,thevine,thecommondogwoodandthecorneliancherry。
TheWhite—girdledLeaf—cutterlikestherobinia,towhichsheadds,inlavishproportions,thevine,theroseandthehawthornandsometimes,inmoderation,thereedandthewhitish—leavedrock—rose。
TheBlack—tippedLeaf—cutter(Megachileapicalis,SPIN。)hasforherabodethecellsoftheMason—beeofthePebblesandtheruinednestsoftheOsmiaeandAnthidiaintheSnail—shells。Ihavenotknownhertouseanyothermaterialsthanthewildbriarandthehawthorn。
Incompletethoughitbe,thislisttellsusthattheMegachilesdonothaveexclusivebotanicaltastes。Eachspeciesmanagesextremelywellwithseveralplantsdifferinggreatlyinappearance。Thefirstconditiontobefulfilledbytheshrubexploitedisthatitbenearthenest。Frugalofhertime,theLeaf—cutterdeclinestogoondistantexpeditions。WheneverIcomeuponarecentMegachile—nest,I
amnotlonginfindingintheneighbourhood,withoutmuchsearching,thetreeorshrubfromwhichtheBeehascutherpieces。
Anothermainconditionisafineandsuppletexture,especiallyforthefirstdisksusedinthelidandforthepieceswhichformtheliningofthewallet。Therest,lesscarefullyexecuted,allowsofcoarserstuff;buteventhenthepiecemustbeflexibleandlenditselftothecylindricalconfigurationofthetunnel。Theleavesoftherock—roses,thickandroughlyfluted,fulfilthisconditionunsatisfactorily,forwhichreasonIseethemoccurringonlyatveryrareintervals。Theinsecthasgatheredpiecesofthembymistakeand,notfindingthemgoodtouse,hasceasedtovisittheunprofitableshrub。Stifferstill,theleafoftheholm—oakinitsfullmaturityisneveremployed:theSilkyLeaf—cutterusesitonlyintheyoungstateandtheninmoderation;shecangethervelvetypiecesbetterfromthevine。Inthelilac—bushessozealouslyexploitedbeforemyeyesbytheHare—footedLeaf—cutteroccuramedleyofdifferentshrubswhich,fromtheirsizeandthelustreoftheirleaves,shouldapparentlysuitthatsturdypinker。Theyaretheshrubbyhare's—ear,thehoneysuckle,thepricklybutcher's—broom,thebox。Whatmagnificentdisksoughttocomefromthehare's—earandthehoneysuckle!Onecouldgetanexcellentpiece,withoutfurtherlabour,bymerelycuttingtheleaf—stalkofthebox,asMegachilesericansdoeswithherpaliurus。Thelilac—loverdisdainsthemabsolutely。Forwhatreason?Ifancythatshefindsthemtoostiff。
Wouldshethinkdifferentlyifthelilac—bushwerenotthere?Perhapsso。
Inshort,apartfromthequestionsoftextureandproximitytothenest,theMegachile'schoice,itseemstome,mustdependuponwhetheraparticularshrubisplentifulornot。Thiswouldexplainthelavishuseofthevine,anobjectofwidespreadcultivation,andofthehawthornandthewildbriar,whichformpartofallourhedges。Asthesearetobefoundeverywhere,thefactthatthedifferentLeaf—cuttersmakeuseofthemisnoreflectionuponahostofequivalentsvaryingaccordingtothelocality。
Ifwehadtobelievewhatpeopletellusabouttheeffectsofheredity,whichissaidtohanddownfromgenerationtogeneration,evermorefirmlyestablished,theindividualhabitsofthosewhocomebefore,theMegachilesoftheseparts,experiencedinthelocalflorabythelongtrainingofthecenturies,butcompletenovicesinthepresenceofplantswhichtheirraceencountersforthefirsttime,oughttorefuseasunusualandsuspiciousanyexoticleaves,especiallywhentheyhaveathandplentyoftheleavesmadefamiliarbyhereditarycustom。Thequestionwasdeservingofseparatestudy。
Twosubjectsofmyobservations,theHare—footedandtheSilveryLeaf—cutter,bothoftheminmatesofmyopen—airlaboratory,gavemeadefiniteanswer。KnowingthepointsfrequentedbythetwoMegachiles,Iplantedintheirwork—yard,overgrownwithbriarandlilac,twooutlandishplantswhichseemedtometofulfiltherequiredconditionsofsupplenessoftexture,namely,theailantus,anativeofJapan,andtheVirginianphysostegia。Eventsjustifiedtheselection:bothBeesexploitedtheforeignflorawiththesameassiduityasthelocalflora,passingfromthelilactotheailantus,fromthebriartothephysostegia,leavingtheone,goingbacktotheother,withoutdrawingdistinctionsbetweentheknownandtheunknown。Inveteratehabitcouldnothavegivengreatercertainty,greatereasetotheirscissors,thoughthiswastheirfirstexperienceofsuchamaterial。
TheSilveryLeaf—cutterlentherselftoanevenmoreconclusivetest。
Asshereadilymakeshernestinthereedsofmyapparatus,Iwasable,uptoacertainpoint,tocreatealandscapeforherandselectitsvegetationmyself。Ithereforemovedthereed—hivetoapartoftheenclosurestockedchieflywithrosemary,whosescantyfoliageisnotadaptedfortheBee'swork,andneartheapparatusIarrangedanexoticshrubberyinpots,includingnotablythesmoothlopezia,fromMexico,andthelong—fruitedcapsicum,anIndianannual。Findingcloseathandthewherewithaltobuildhernest,theLeaf—cutterwentnofurtherafield。Thelopeziasuitedherespecially,somuchsothatalmostthewholenestwascomposedofit。Theresthadbeengatheredfromthecapsicum。
Anotherrecruit,whoseco—operationIhadinnowayengineered,camespontaneouslytooffermeherevidence。ThiswastheFeebleLeaf—
cutter(Megachileimbecilla,GERST。)。Nearlyaquarterofacenturyago,Isawher,allthroughthemonthofJuly,cuttingoutherroundsandellipsesattheexpenseofthepetalsofthePelargoniumzonale,thecommongeranium。Herperseverancedevastated——thereisnootherwordforit——mymodestarrayofpots。Hardlywasablossomout,whentheardentMegachilescameandscallopeditintocrescents。Thecolourwasindifferenttoher:red,whiteorpink,allthepetalsunderwentthedisastrousoperation。Afewcaptures,ancientrelicsofmycollecting—boxesbythistime,indemnifiedmeforthepillage。I
havenotseenthisunpleasantBeesince。Withwhatdoesshebuildwhentherearenogeranium—flowershandy?Idonotknow;butthefactremainsthatthefragiletailoressusedtoattacktheforeignflower,afairlyrecentacquisitionfromtheCape,asthoughallherracehadneverdoneanythingelse。
Thesedetailsleaveuswithoneobviousconclusion,whichiscontrarytoouroriginalideas,basedontheunvaryingcharacterofinsectindustry。Inconstructingtheirjars,theLeaf—cutters,eachfollowingthetastepeculiartoherspecies,donotmakeuseofthisorthatplanttotheexclusionoftheothers;theyhavenodefiniteflora,nodomainfaithfullytransmittedbyheredity。Theirpiecesofleavesvaryaccordingtothesurroundingvegetation;theyvaryindifferentlayersofthesamecell。Everythingsuitsthem,exoticornative,rareorcommon,providedthatthebitcutoutbeeasytoemploy。Itisnotthegeneralaspectoftheshrub,withitsfragileorbushybranches,itslargeorsmall,greenorgrey,dullorglossyleaves,thatguidestheinsect:suchadvancedbotanicalknowledgedoesnotenterintothequestionatall。Inthethicketchosenasapinking—establishment,theMegachileseesbutonething:leavesusefulforherwork。TheShrike,withhispassionforplantswithlong,woollysprigs,knowswheretofindnicely—waddedsubstituteswhenhisfavouritegrowth,thecotton—rose,islacking;theMegachilehasmuchwiderresources:indifferenttotheplantitself,shelooksonlyintothefoliage。Ifshefindsleavesofthepropersize,ofadrytexturecapableofdefyingthedampandofasupplenessfavourabletocylindricalcurving,thatisallsheasks;andtherestdoesnotmatter。Shehasthereforeanalmostunlimitedfieldforherlabour。
Thesesuddenandwhollyunprovokedchangesgivecauseforreflection。
Whenmygeranium—flowersweredevastated,howhadtheobtrusiveBee,untroubledbytheprofounddissimilaritybetweenthepetals,snow—
whitehere,brightscarletthere,howhadshelearnthertrade?
NothingtellsusthatsheherselfwasnotforthefirsttimeexploitingtheplantfromtheCape;and,ifshereallydidhavepredecessors,thehabithadnothadtimetobecomeinveterate,consideringthemodernimportationofthegeranium。WhereagaindidtheSilveryMegachile,forwhomIcreatedanexoticshrubbery,maketheacquaintanceofthelopezia,whichcomesfromMexico?Shecertainlyismakingafirststart。Neverdidhervillageorminepossessastalkofthatchillydenizenofourhot—houses。Sheismakingafirststart;andbeholdherstraightwayagraduate,versedintheartofcarvingunfamiliarfoliage。
Peopleoftentalkofthelongapprenticeshipsservedbyinstinct,ofitsgradualacquirements,ofitstalents,thelaboriousworkoftheages。TheMegachilesaffirmtheexactopposite。Theytellmethattheanimal,thoughinvariableintheessenceofitsart,iscapableofinnovationinthedetails;butatthesametimetheyassuremethatanysuchinnovationissuddenandnotgradual。Nothingpreparestheinnovations,nothingimprovesthemorhandsthemdown;otherwiseaselectionwouldlongagohavebeenmadeamidthediversityoffoliage;andtheshrubrecognizedasthemostserviceable,especiallywhenitisalsoplentiful,wouldalonesupplyallthebuilding—
materialsneeded。Ifhereditytransmittedindustrialdiscoveries,aMegachilewhothoughtofcuttingherdisksoutofpomegranate—leavesandfoundthemsatisfactoryoughttohaveinstilledalikingforsimilarmaterialsintoherdescendants;andweshouldthisdayfindLeaf—cuttersfaithfultothepomegranate—leaves,workerswhoremainedexclusiveintheirchoiceoftherawmaterial。Thefactsrefutethesetheories。
Peoplealsosay:
'Grantusavariation,howeversmall,intheinsect'sindustry;andthatvariation,accentuatedmoreandmore,willproduceanewraceandfinallyafixedspecies。'
ThistriflingvariationisthefulcrumforwhichArchimedesclamouredinordertolifttheworldwithhissystemoflevers。TheMegachilesofferusoneandaverygreatone:theindefinitevariationoftheirmaterials。Whatwillthetheorists'leversliftwiththisfulcrum?
Why,nothingatall!Whethertheycutthedelicatepetalsofthegeraniumorthetoughleavesofthelilac—bushes,theLeaf—cuttersareandwillbewhattheywere。Thisiswhatwelearnfromthepersistenceofeachspeciesinitsstructuraldetails,despitethegreatvarietyofthefoliageemployed。
CHAPTER9。THECOTTON—BEES。
TheevidenceoftheLeaf—cuttersprovesthatacertainlatitudeislefttotheinsectinitschoiceofmaterialsforthenest;andthisisconfirmedbythetestimonyoftheAnthidia,thecotton—
manufacturers。Mydistrictpossessesfive:A。Florentinum,LATR。,A。
diadema,LATR。,A。manicatum,LATR。,A。cingulatum,LATR。,A。
scapulare,LATR。Noneofthemcreatestherefugeinwhichthecottongoodsaremanufactured。LiketheOsmiaeandtheLeaf—cutters,theyarehomelessvagrants,adopting,eachtoherowntaste,suchshelterastheworkofothersaffords。TheScapularAnthidiumisloyaltothedrybramble,deprivedofitspithandturnedintoahollowtubebytheindustryofvariousminingBees,amongwhichfigure,inthefrontrank,theCeratinae,dwarfrivalsoftheXylocopa,orCarpenter—bee,thatmightydrillerofrottenwood。ThespaciousgalleriesoftheMaskedAnthophorasuittheFlorentineAnthidium,theforemostmemberofthegenussofarassizeisconcerned。TheDiademAnthidiumconsidersthatshehasdoneverywellifsheinheritsthevestibuleoftheHairy—footedAnthophora,oreventheordinaryburrowoftheEarth—worm。Failinganythingbetter,shemayestablishherselfinthedilapidateddomeoftheMason—beeofthePebbles。TheManicateAnthidiumshareshertastes。IhavesurprisedtheGirdledAnthidiumcohabitingwithaBembex—wasp。Thetwooccupantsofthecaveduginthesand,theownerandthestranger,werelivinginpeace,bothintentupontheirbusiness。Herusualhabitationissomeholeorotherinthecrevicesofaruinedwall。Totheserefuges,theworkofothers,wecanaddthestumpsofreeds,whichareaspopularwiththevariouscotton—gatherersaswiththeOsmiae;and,afterwehavementionedafewmostunexpectedretreats,suchasthesheathprovidedbyahollowbrickorthelabyrinthfurnishedbythelockofagate,weshallhavealmostexhaustedthelistofdomiciles。
LiketheOsmiaeandtheLeaf—cutters,theAnthidiumshowsanurgentneedofaready—madehome。Sheneverhousesherselfatherownexpense。Canwediscoverthereason?Letusfirstconsultafewhardworkerswhoareartificersoftheirowndwellings。TheAnthophoradigscorridorsandcellsintheroad—sidebankshardenedbythesun;
shedoesnoterect,sheexcavates;shedoesnotbuild,sheclears。
Toilingawaywithhermandibles,atombyatom,shemanagestocontrivethepassagesandchambersnecessaryforhereggs;andahugebusinessitis。Shehas,inaddition,topolishandglazetheroughsidesofhertunnels。Whatwouldhappenif,afterobtainingahomebydintoflong—continuedtoil,shehadnexttolineitwithwadding,togatherthefibrousdownfromcottonyplantsandtofeltitintobagssuitableforthehoney—paste?Thehard—workingBeewouldnotbeequaltoproducingalltheserefinements。Herminingcallsfortoogreatanexpenditureoftimeandstrengthtoleavehertheleisureforluxuriousfurnishing。Chambersandcorridors,therefore,willremainbare。
TheCarpenter—beegivesusthesameanswer。Whenwithherjoiner'swimbleshehaspatientlyboredthebeamtoadepthofnineinches,wouldshebeabletocutoutandplaceinpositionthethousandandonepieceswhichtheSilkyLeaf—cutteremploysforhernest?Timewouldfailher,evenasitwouldfailaMegachilewho,lackingtheCapricorn'schamber,hadherselftodigahomeinthetrunkoftheoak。ThereforetheCarpenter—bee,afterthetediousworkofboring,getstheinstallationdoneinthemostsummaryfashion,simplyrunningupasawdustpartition。
Thetwothings,thelaboriousbusinessofobtainingalodgingandtheartisticworkoffurnishing,seemunabletogotogether。Withtheinsectaswithman,hewhobuildsthehousedoesnotfurnishit,hewhofurnishesitdoesnotbuildit。Toeachhisshare,becauseoflackoftime。Divisionoflabour,themotherofthearts,makestheworkmanexcelinhisdepartment;onemanforthewholeworkwouldmeanstagnation,theworkernevergettingbeyondhisfirstcrudeattempts。Animalindustryisalittlelikeourown:itdoesnotattainitsperfectionsavewiththeaidofobscuretoilers,who,withoutknowingit,preparethefinalmasterpiece。IseenootherreasonforthisneedofagratuitouslodgingfortheMegachile'sleafybasketortheAnthidia'scottonpurses。Inthecaseofotherartistswhohandledelicatethingsthatrequireprotection,Idonothesitatetoassumetheexistenceofaready—madehome。ThusReaumurtellsusoftheUpholsterer—bee,Anthocopapapaveris,whofashionshercellswithpoppy—petals。Idonotknowtheflower—cutter,Ihaveneverseenher;butherarttellsmeplainlyenoughthatshemustestablishherselfinsomegallerywroughtbyothers,as,forinstance,inanEarth—worm'sburrow。
WehavebuttoseethenestofaCotton—beetoconvinceourselvesthatitsbuildercannotatthesametimebeanindefatigablenavvy。
Whenandnewly—feltedandnotyetmadestickywithhoney,thewaddedpurseisbyfarthemostelegantknownspecimenofentomologicalnest—building,especiallywherethecottonisofabrilliantwhite,asisfrequentlythecaseinthemanufacturersoftheGirdledAnthidium。Nobird's—nest,howeverdeservingofouradmiration,canvieinfinenessofflock,ingracefulnessofform,indelicacyoffeltingwiththiswonderfulbag,whichourfingers,evenwiththeaidoftools,couldhardlyimitate,foralltheirdexterity。Iabandontheattempttounderstandhow,withitslittlebalesofcottonbroughtuponebyone,theinsect,nootherwisegiftedthanthekneadersofmudandthemakersofleafybaskets,managestofeltwhatithascollectedintoahomogeneouswholeandthentoworktheproductintoathimble—shapedwallet。Itstoolsasamaster—fullerareitslegsanditsmandibles,whicharejustlikethosepossessedbythemortar—kneadersandLeaf—cutters;andyet,despitethissimilarityofoutfit,whatavastdifferenceintheresultsobtained!
ToseetheCotton—bees'talentsinactionseemsanundertakingfraughtwithinnumerabledifficulties:thingshappenatadepthinaccessibletotheeye;andtopersuadetheinsecttoworkintheopendoesnotlieinourpower。OneresourceremainedandIdidnotfailtoturntoit,thoughhithertoIhavebeenwhollyunsuccessful。
Threespecies,Anthidiumdiadema,A。manicatumandA。florentinum——
thefirst—namedinparticular——showthemselvesquitereadytotakeuptheirabodeinmyreed—apparatus。AllthatIhadtodowastoreplacethereedsbyglasstubes,whichwouldallowmetowatchtheworkwithoutdisturbingtheinsect。ThisstratagemhadansweredperfectlywiththeThree—hornedOsmiaandLatreille'sOsmia,whoselittlehousekeeping—secretsIhadlearntthankstothetransparentdwelling—
house。WhyshoulditnotanswerforitsCotton—beesand,inthesameway,withtheLeaf—cutters?Ialmostcountedonsuccess。Eventsbetrayedmyconfidence。ForfouryearsIsuppliedmyhiveswithglasstubesandnotoncedidtheCotton—weaversortheLeaf—cutterscondescendtotakeuptheirquartersinthecrystalpalaces。Theyalwayspreferredthehovelprovidedbythereed。ShallIpersuadethemoneday?Idonotabandonallhope。
Meanwhile,letmedescribethelittlethatIsaw。Moreorlessstockedwithcells,thereedisatlastclosed,rightattheorifice,withathickplugofcotton,usuallycoarserthanthewaddingofthehoney—satchels。ItistheequivalentoftheThree—hornedOsmia'sbarricadeofmud,oftheleaf—puttyofLatreille'sOsmia,oftheMegachiles'barrierofleavescutintodisks。Allthesefreetenantsarecarefultoshuttightthedoorofthedwelling,ofwhichtheyhaveoftenutilizedonlyaportion。Towatchthebuildingofthisbarricade,whichisalmostexternalwork,demandsbutalittlepatienceinwaitingforthefavourablemoment。
TheAnthidiumarrivesatlast,carryingthebaleofcottonfortheplugging。Withherfore—legsshetearsitapartandspreadsitout;
withhermandibles,whichgoinclosedandcomeoutopen,sheloosensthehardlumpsofflock;withherforeheadshepresseseachnewlayerupontheonebelow。Andthatisall。Theinsectfliesoff,returnsthericherbyanotherbaleandrepeatstheperformanceuntilthecottonbarrierreachestheleveloftheopening。Wehavehere,remember,aroughtask,innowaytobecomparedwiththedelicatemanufacturerofthebags;nevertheless,itmayperhapstellussomethingofthegeneralprocedureofthefinerwork。Thelegsdothecarding,themandiblesthedividing,theforeheadthepressing;andtheplayoftheseimplementsproducesthewonderfulcushionedwallet。
Thatisthemechanisminthelump;butwhatoftheartistry?
Letusleavetheunknownforfactswithinthescopeofobservation。I
willquestiontheDiademAnthidiuminparticular,afrequentinmateofmyreeds。Iopenareed—stumpabouttwodecimetreslongbytwelvemillimetresindiameter。(Aboutsevenandthree—quarterinchesbyhalfaninch。——Translator'sNote。)Theendisoccupiedbyacolumnofcotton—woolcomprisingtencells,withoutanydemarcationbetweenthemontheoutside,sothattheirwholeformsacontinuouscylinder。
Moreover,thankstoaclosefelting,thedifferentcompartmentsaresolderedtogether,somuchsothat,whenpulledbytheend,thecottonedificedoesnotbreakintosections,butcomesoutallinonepiece。Onewouldtakeitforasinglecylinder,whereasinrealitytheworkiscomposedofaseriesofchambers,eachofwhichhasbeenconstructedseparately,independentlyoftheonebefore,exceptperhapsatthebase。
Forthisreason,shortofrippingupthesoftdwelling,stillfullofhoney,itisimpossibletoascertainthenumberofstoreys;wemustwaituntilthecocoonsarewoven。Thenourfingerscantellthecellsbycountingtheknotsthatresistpressureunderthecoverofwadding。Thisgeneralstructureiseasilyexplained。Acottonbagismade,withthesheathofthereedasamould。Ifthisguidingsheathwerelacking,thethimbleshapewouldbeobtainedallthesame,withnolesselegance,asisprovedbytheGirdledAnthidium,whomakeshernestinsomehiding—placeorotherinthewallsortheground。
Whenthepurseisfinished,theprovisionscomeandtheegg,followedbytheclosingofthecell。WedonotherefindthegeometricallidoftheLeaf—cutters,thepileofdiskstight—setinthemouthofthejar。Thebagisclosedwithacottonsheetwhoseedgesaresolderedbyafelting—processtotheedgesoftheopening。Thesolderingissowelldonethatthehoney—pouchanditscoverformanindivisiblewhole。Immediatelyaboveit,thesecondcellisconstructed,havingitsownbase。Atthebeginningofthiswork,theinsecttakescaretojointhetwostoreysbyfeltingtheceilingofthefirsttothefloorofthesecond。Thuscontinuedtotheend,thework,withitsinnersolderings,becomesanunbrokencylinder,inwhichthebeautiesoftheseparatewalletsdisappearfromview。Inverymuchthesamefashion,butwithlessadhesionamongthedifferentcells,dotheLeaf—cuttersactwhenstackingtheirjarsinacolumnwithoutanyexternaldivisionintostoreys。